An organism’s appearance is the result of evolutionary pressures, and those same pressures apply to the structures organisms build, such as nests. Superorganism nests function as extended phenotypes to perform key biological processes, to survive, grow, and reproduce. Social insects are masters of solving organizational problems because they must coordinate thousands of individuals to accomplish these goals. One such problem is how to construct nests, and then, how to organize resources within that nest. Both, presumably, are optimized to maximize colony performance.