PhD in Physics, University of Chicago, 1968 (thesis: Fast Photovoltages from Photoreceptors).
Post doc, 1967-68, with Rod Clayton at Cornell (Photosynthesis of Bacteria)
Assistant Professor, 1968-73, Columbia University, Dept of Biological Sciences
Associate Prof--Prof, 1973-2000, University of Illinois at Urbana-Champaign, Depts of Physiology and Biophysics, and Cell and Structural Biology
Research Professor, University of Washington, Dept of Biology, 2000-