Resident Assoc. Director of the Friday Harbor Labs, UW, from 1973 to 2008. Also since 1973, on the faculty of the Zoology Department and now Biology Department at the UW to 2008.
Previously, a Predoctoral Fellow, NSF, Univ. of Washington 1963-69; Postdoctoral Trainee, NIH, UCLA, 1970 Postdoctoral Fellow, NSF, Univ. of Hawaii, 1970-71 Assistant Prof. in Dept. of Zoology, Univ. of Maryland, 1971-73. NSF grants: Regional patterns in reproduction and survival of the barnacle Balanus glandula (1975-1981), Larvae feeding with cilia (1981-83), Constraints on foraging and form associated with two larval ciliary feeding mechanisms (1984-86), The relation between adult size, brooding, and simultaneous hermaphroditism (with D. Eernisse) (1984-86), Constraints on modes of development of benthic marine invertebrates (1986-1990), Interocean comparisons of food limited larval growth (1987-1988), Limits on size and form of benthic clutches (1990-1993), Plasticity of larval feeding structures (1993-1996), Duration of development of planktonic embryos (1996-2001), Constraints on performance and pelagic periods of larvae (2001-2004), Swimming and vulnerability in pelagic development (2002-2007), Predator-induced morphological defenses in marine zooplankton (2006-2009). Guggenheim Fellow for research on sea urchin form at Smithsonian Inst. and on ciliary feeding at Marine Biol. Assoc. UK, 1980-81