Professor Emeritus
206-543-4066 (office)

Research Overview

My science is inspired by observations that appear to be a challenge to explain by natural selection. My early work here focused on field experimental investigations of the evolution of badge signaling systems and how they resist invasion by cheaters, on parent cannibalism of offspring, and on the evolution of adoption of unrelated offspring by replacement mates. Later, as we developed a premier bird collection for the Northwest at the Burke Museum, my interests expanded to include a diversity of projects that are well suited to investigations using museum collections.


I grew up in the deep south where I spent most of my childhood in the woods catching snakes. Glen Woolfenden was my undergraduate mentor and the only field biologist at the University of South Florida in the 60