Professor Emeritus
206-543-1976 (office)

Research Overview

Ecology and Evolution: (1) Adaptations of plants to azonal (unusual) soils, especially serpentines (2) Nickel hyperaccumulation in serpentine plants (3) Evolution of tolerance to serpentine soils.

Systematics and Taxonomy: (1) Speciation in serpentine endemic plants (2) Taxonomy of Streptanthus (Section Euclisia, Cruciferae) (3) Interfertility among North American species of Silene and Lychnis (Caryophyllaceae) (4) Cytogenetics of interspecific hybrids (5) Cytotaxonomy of Silene and Phacelia.